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Las Cerradicas is located towards the top of the Villar del Arzobispo Formation (early Berriasian). This location adds to the significance of the dinosaur track record of the Parque Nacional Torotoro (Torotoro National Park), Bolivia.Ī study of the undescribed part of the Las Cerradicas tracksite in Galve (Teruel, Spain) has revealed 6 trackways made by small sauropods showing gregarious behaviour (parallel trackways). This report records abundant theropod tracks and swim traces in a fortuitous exposure that is no longer accessible and comments on behavioral and paleoenvironmental implications. Before substrate stabilization, the water level rose, and dinosaurs swam in the area, leaving characteristic swim trackways with their feet scratching the bottom. Invertebrate burrowing and sediment collapse, after the kick-off phase of track formation, partially modified most tracks.

Swim traces occurred in three distinct morphotypes. We distinguished five track morphotypes based on the morphology of the digits and pedal impression. In addition to tracks, 73 swim traces from 14 trackways were characterized. Of those trackways, 21 consisted of a single, isolated track. The upper bed contained 493 tracks in 88 trackways. Due to time constraints, only the upper layer containing a rich assemblage of theropod tracks and traces was thoroughly studied before road construction removed the bed. In the lower layer, five theropod and five sauropod trackways were identified, three trackways of uncertain identity, and three trackways of didactyl tracks. The trackways were found on two beds of ooid and ostracod-bearing quartz arenites of about 160 m 2, each separated by roughly 2 m of calcareous siltstone. , a new site containing dinosaur trackways was uncovered at kilometer 99 during the Anzaldo to Torotoro highway construction.

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